Priscila Chaverri Echandi, Ph.D.

Universidad de Costa Rica

Escuela de Biología

Apdo. 11501-2060

San Pedro, San José


Correo electrónico:




(*Estudiantes de posgrado y post-doctorado bajo supervisión o tutorías de P. Chaverri)


Salgado-Salazar, C.*, Rossman, A.Y., Chaverri, P. 2016. The genus Thelonectria (Nectriaceae, Hypocreales, Ascomycota) and closely related species with cylindrocarpon-like asexual states. Fungal Diversity (en revisión)

Hirooka, Y.*, Rossman, A.Y., Samuels, G.J., Lechat, C., Chaverri, P. 2012. A monograph of Allantonectria, Nectria, and Pleonectria (Nectriaceae, Hypocreales, Ascomycota) and their pycnidial, sporodochial, and synnematous anamorphs. Studies in Mycology 71: 1-210. (Open Access)

Chaverri, P., Huhndorf, S., Rogers, J., Samuels, G.J. 2010. Microhongos Comunes de Costa Rica y otras regiones tropicales (Ascomycota, Pezizomycotina, Sordariomycetes). Common Microfungi of Costa Rica and other tropical regions (Ascomycota, Pezizomycotina, Sordariomycetes). INBio Editorial: Heredia. 241 pp. + 60 Pl.

Chaverri, P., Liu, M., Hodge, K.T. 2008. Neotropical Hypocrella (anamorph Aschersonia), Moelleriella, and Samuelsia. Studies in Mycology 60: 1-68. (Open Access)

Samuels, G.J., Rossman, A.Y., Chaverri, P., Overton, BE, Põldmaa, K. 2006. Hypocreales of the Southeastern United States: An Identification Guide. CBS Biodiversity Series No. 4. Centraalbureau voor Schimmelcultures (CBS): Utrecht. 145 pp. (

Chaverri, P., Samuels, G.J. 2003. Hypocrea/Trichoderma (Ascomycota, Hypocreales, Hypocreaceae): Species with green ascospores. Studies in Mycology 48: 1-116 Trichoderma Clave Interactiva.



Vidaurre Montoya, Q., Andrade Meirelles, L., Chaverri, P., Rodrigues, A. 2016. Unraveling Trichoderma species in attine ant environment: description of three new taxa. Antonie van Leeuwenhoek Journal of Microbiology (aceptado).

Herrera, C.S.*, Hirooka, Y., Chaverri, P. 2016. Pseudocospeciation of the mycoparasite Cosmospora with their associated fungal hosts. Ecology and Evolution doi:10.1002/ece3.1967.

Gazis, R.*, Chaverri, P. 2015. Wild trees in the Amazon basin harbor a great diversity of beneficial endosymbiotic fungi: Is this evidence of protective mutualism? Fungal Ecology 17: 18-29.

Rosmana, A., Samuels, G.J., Ismaiel, A., Ibrahim, E.S., Chaverri, P., Herawati, Y. 2015. Trichoderma asperellum: A dominant endophyte species in cacao grown in Sulawesi with potential for controlling Vascular Streak Dieback disease. Tropical Plant Pathology (en imprenta). doi 10-1007/s40858-015-0004-1.

Herrera, C.S.*, Rossman, A.Y., Samuels, G.J., Liparini-Pereira, O., Chaverri, P. 2015. Revision of the Cosmospora viliuscula species complex. Mycologia (en imprenta).

Chaverri, P., Branco-Rocha, F., Jaklitsch, W., Degenkolb, T., Gazis, R.*, Samuels, G.J. 2015. Systematics of the Trichoderma harzianum species complex and the reidentification of commercial biocontrol strains. Mycologia (en imprenta) doi 10.3852/14-147.

Degenkolb T., Nielsen K.F., Dieckmann R., Branco-Rocha F., Chaverri P., Samuels G.J., Thrane U., von Döhren H., Vilcinskas A., Brückner H. 2014[2015]. Peptaibol, secondary metabolite, and hydrophobin pattern of commercial biocontrol agents formulated with species of the Trichoderma harzianum complex. Chemistry and Biodiversity (en imprenta) doi 10.1002/cbdv.201400300.

Martin, R., Gazis, R., Skaltsas, D.*, Chaverri, P., Hibbett, D. 2015. Unexpected diversity of basidiomycetous endophytes in sapwood and leaves of Hevea. Mycologia (en imprenta). doi:10.3852/14-206

Gazis, R.*, Skaltsas, D.*, Chaverri, P. 2014. Novel endophytic lineages of Tolypocladium provide new insights into the ecology and evolution of Cordyceps-like fungi. Mycologia 106: 1090-1105.

Salgado-Salazar, C.,* Rossman, A.Y., Samuels, G.J., Hirooka, Y., Sanchez, R.M., Chaverri, P. 2014. Phylogeny and taxonomic revision of Thelonectria discophora (Ascomycota, Hypocreales, Nectriaceae) species complex. Fungal Diversity (en imprenta).

Salgado-Salazar, C.*, Rossman, A.Y., Chaverri, P. 2013. Not as ubiquitous as we thought: taxonomic crypsis, hidden diversity and cryptic speciation in the cosmopolitan fungus Thelonectria discophora (Nectriaceae, Hypocreales, Ascomycota). PLoS ONE 8(10): e76737. doi:10.1371/journal.pone.0076737

Chaverri, P., Samuels, G.J. 2013. Evolution of habitat preference and nutrition mode in a cosmopolitan fungal genus with evidence of interkingdom host jumps and major shifts in ecology. Evolution 67: 2823–2837. PDF

Herrera, C.S.*, Rossman, A.Y., Samuels, G.J., Chaverri, P. 2013. Pseudocosmospora - a new genus to accommodate Cosmospora vilior and related species. Mycologia 105: 1287-1305.

Herrera, C.S.*, Rossman, A.Y., Samuels, G.J. Lechat, C., Chaverri, P. 2013. Revision of the genus Corallomycetella with Corallonectria gen. nov. for C. jatrophae (Nectriaceae, Hypocreales). Mycosystema 32: 518-544 (edición especial dedicada a R.F. Korf). PDF

Hirooka, Y.*, Rossman, A.Y., Zhuang, W-Y., Salgado, C., Chaverri, P. 2013. Species delimitation for Neonectria coccinea group including the causal agents of beech bark disease (BBD) in Asia, Europe, and North America. Mycosystema 32: 485-517 (edición especial dedicada a R.F. Korf). PDF

Unterseher, M., Gazis, R.*, Chaverri, P., García Guarniz, C.F., Zavaleta Tenorio, D.H. 2013. Endophytic fungi from Peruvian highland and lowland habitats form distinctive and host plant specific assemblages. Biodiversity and Conservation 22: 999-1016. PDF

Rossman, A.Y., Seifert, K.A., Samuel, G.J., Minnis, A.M., Schroers, H.-J., Lombard, L., Crous, P.W., Põldma, K., Cannon, P.F., Summerbel, R.C., Geiser, D.M., Zhuan, W.-Y., Hirooka, Y.*, Herrera, C.*, Salgado-Salazar, C.*, Chaverri, P. 2013. Genera in Bionectriaceae, Hypocreaceae, and Nectriaceae (Hypocreales) proposed for acceptance or rejection. IMA Fungus 4: 41-51. PDF

Gazis, R.*, Miadlikowska, J., Lutzoni, F., Arnold, A.E., Chaverri, P. 2012. Rubber trees in Peru host a new class of fungi: ‘Xylonomycetes’ (Ascomycota). Molecular Phylogenetics and Evolution 65: 294-304. PDF

Salgado, C.*, Rossman, A.Y, Samuels, G.J., M. Capdet, Chaverri, P. 2012. Multigene phylogenetic analyses of the Thelonectria coronata and T. veuillotiana species complexes. Mycologia 104: 1325-1350. PDF

Schoch, C.L., Seifert, K.A., Huhndorf, S., Robert, V., Spouge, J.L., Levesque, A.,Chen, W., and Fungal Barcoding Consortium**. 2012. Nuclear ribosomal internal transcribed spacer (ITS) region as a universal DNA barcode marker for Fungi. Proceedings of the National Academy of Sciences 109: 6241-6246. **P. Chaverri es parte del "Fungal Barcoding Consortium" o Consorcio para Código de Barras de Hongos. PDF

Samuels, G.J., Ismaiel, A., de Souza, J., Chaverri, P. 2012. Trichoderma stromaticum and its overseas relatives. Mycological Progress 11: 215-254. PDF

Gazis, R.*, Rehner, S., Chaverri, P. 2011. Species delimitation in fungal endophyte diversity studies and its implications in ecological and biogeographic inferences. Molecular Ecology 20: 3001-3013. PDF. (Artículo destacado en la sección PERSPECTIVES de Mol. Ecol., 2011, 20: 2873-2875)

Hirooka, Y.*, Rossman, A.Y., Chaverri, P. 2011. Morphological and phylogenetic analyses of the Nectria cinnabarina species complex. Studies in Mycology 68: 35-56. Open Access.

Chaverri, P., Salgado, C.*, Hirooka, Y.*, Rossman, A.Y., Samuels, G.J. 2011. Delimitation of Neonectria and Cylindrocarpon and related genera with Cylindrocarpon-like anamorphs. Studies in Mycology 68: 57-78. Open Access.

Chaverri, P., Gazis, R.*, Samuels, G.J. 2011. Trichoderma amazonicum, a new endophytic species on Hevea brasiliensis and H. guianensis from the Amazon basin. Mycologia 103: 139-151. PDF

Chaverri, P., Gazis, R.* 2011. Linking ex planta fungi with their endophytic stages: Perisporiopsis, a common leaf litter and soil fungus, is a frequent endophyte of Hevea spp. and other plants. Fungal Ecology 4: 94-102. PDF

Chaverri, P., Gazis, R.* 2010. Perisporiopsis lateritia, a new species on decaying leaves of Hevea spp. from the Amazon basin in Peru. Mycotaxon 113: 163-169. PDF

Gazis R.*, Chaverri, P.2010. Diversity of fungal endophytes in leaves and stems of wild rubber trees (Hevea brasiliensis) in Peru. Fungal Ecology 3: 240-254. PDF

Hirooka, Y.*, Kobayashi, T., Ono, T., Rossman, A.Y., Chaverri, P. 2010. Verrucostoma, a new genus in the Bionectriaceae from the Bonin Islands, Japan. Mycologia 102: 418-429. PDF

Minnis, D., Samuels, G.J., Chaverri, P. 2009. Three nomenclatural corrections for species of Hypocrea/Trichoderma. Mycotaxon 109: 245-246.

Liu, M., Milgroom, M.G., Chaverri, P., Hodge, K.T. 2009. Speciation of a tropical fungal species pair following transoceanic dispersal. Molecular Phylogenetics and Evolution 51: 413-426. PDF

Tadych, M., Chaverri, P., Bergen, M., White, J.F. 2009. Moelleriella zhongdongii: stromal development and identification of Hirsutella-like and Aschersonia synanamorphs. Mycological Research 113: 611-615. PDF

Samuels, G.J., Lu, B.-S., Chaverri, P., Candoussau, F., Fournier, J., Rossman, A.Y. 2009. Cyanonectria, a new genus for Nectria cyanostoma and its Fusarium anamorph. Mycological Progress 8: 49-58. PDF

Rojas, E.I., Herre, E.A., Mejía, L.C., Arnold, A.E., Chaverri, P., Samuels, G.J. 2008. Endomelanconiopsis endophyticum, a new Botryosphaeria leaf endophyte from Panama. Mycologia 100: 760-775. PDF

Degenkolb, T., Ralf, D., Nielsen, K.F., Grafenhan, T., Theis, C., Zafari, D., Chaverri, P., Ismaiel, A., Bruckner, H., Von Dohren, H., Thrane, U., Petrini, O., Samuels, G.J. 2008. The Trichoderma brevicompactum clade: a new lineage with new species, new peptaibiotics, and mycotoxins. Mycological Progress 7: 177-209. PDF

Chaverri, P., Vílchez, B. 2006. Hypocrealean (Hypocreales, Ascomycota) fungal diversity in different stages of succession in a tropical forest in Costa Rica. Biotropica 38: 531-543. PDF

Liu, M., Chaverri, P., Hodge, K.T. 2006. A taxonomic revision of the insect biocontrol fungus Aschersonia aleyrodis, its allies with white stromata and their Hypocrella sexual states. Mycological Research 110: 537-554. PDF

Chaverri, P., Bischoff, J.F., Evans, H.C., Hodge, K.T. 2005. Regiocrella, a new entomopathogenic genus with a pycnidial anamorph and its phylogenetic placement in the Clavicipitaceae. Mycologia 97: 1225-1237. PDF

Chaverri, P., Bischoff, J.F., Hodge, K.T. 2005. A new species of Hypocrella, H. macrostroma, and its relationship to other species with large stromata. Mycological Research 109: 1268-1275. PDF

Bischoff, J.F., Chaverri, P., White, J.F. 2005. Clarification of the host substrate of Ascopolyporus (Clavicipitaceae, Hypocreales) and description of Ascopolyporus philodendrus sp. nov. Mycologia 97: 710-717. PDF

Chaverri, P., Samuels, G. J., Hodge, K. T. 2005. The genus Podocrella and its nematode-killing anamorph Harposporium. Mycologia 97: 433-443. PDF

Druzhinina, I. S., Chaverri, P., Fallah, P., Kubicek, C. P., Samuels, G. J. 2004. Hypocrea flaviconidia, a new species from Costa Rica with yellow conidia. Studies in Mycology 50: 401-407. PDF

Chaverri, P., Candoussau, F., Samuels, G.J. 2004. Hypocrea phyllostachydis and its Trichoderma anamorph, a new bambusicolous species from France. Mycological Progress 3: 29-36. PDF

Arguedas, M., Chaverri, P., Verjans, J-M. 2004. Problemas fitosanitarios de la teca en Costa Rica. Revista Recursos Naturales y Ambiente 41: 130-135. PDF

Lu, B.S., Druzhinina, I.S., Fallah, P., Chaverri, P., Gradinger, C., Kubicek, C.P., Samuels, G.J. 2004. Hypocrea/Trichoderma species with pachybasium-like conidiophores: teleomorphs for T. minutisporum and T. polysporum, and their newly discovered relatives. Mycologia 96: 310-342. PDF

Chaverri, P., Castlebury, L.A., Overton, B.E., Samuels, G.J. 2003. Hypocrea/Trichoderma: species with conidiophore elongations and green conidia. Mycologia 95: 1100-1140. PDF (this is a large file)

Chaverri, P., Castlebury, L.A., Samuels, G.J., Geiser, D.M. 2003. Multilocus phylogenetic structure of Trichoderma harzianum/Hypocrea lixii complex. Molecular Phylogenetics and Evolution 27: 302-313. PDF

Dodd, S.L., Lieckfeldt, E., Chaverri, P., Overton, B.E., Samuels, G.J. 2002. Taxonomy and phylogenetic relationships of two species of Hypocrea with Trichoderma anamorphs. Mycological Progress 1: 409-428. PDF

Chaverri, P., Samuels, G.J. 2002. Hypocrea lixii, the teleomorph of Trichoderma harzianum. Mycological Progress 1: 283-286. PDF

Chaverri, P., Samuels, G.J., Stewart, E.L. 2001. Hypocrea virens sp. nov., the teleomorph of Trichoderma virens. Mycologia 93: 1113-1124. PDF

Chaverri, P., Samuels, G.J., Stewart, E.L., Umaña, L. 2001. Hypocrea nigrovirens sp. nov., a new species with a gliocladium-like anamorph. Mycologia 93:758-763. PDF

Arguedas, M., Chaverri, P. 1999. Problemas fitosanitarios en laurel (Cordia alliodora (Ruiz & Pavón) Oken) en Costa Rica. Revista Tecnología en Marcha 13: 18-24.